While you may be right about CO2 to some degree, there's no question that light is the prime limiting factor to plant growth.
Very true up to a point Neale. If I was to start up a planted tank in a dark cupboard nothing would grow. Adding light would be the trigger for growth. All plants will have a minimum threshold for light, and it will almost certainly vary from plant to plant.
What I mean by "bright light" is not necessarily vast amounts of light, but rather the difference between those plants that are shade tolerant (like Java ferns) and those that are not (like Rotala spp.).
I have grown Rotala rotundifolia, sp “green” and sp “Nanjenshan”. Being fast growing stems, they will always reach for the light, meaning that they will rarely spend a great deal of time in the shade in a planted tank.
On the subject of the Rotala sp reaching for the light, there is food for thought that they may be reaching to the higher levels of CO2 that occur higher up in the water column. Hemianthus micranthemoides can sometimes creep along the substrate rather than grow upwards. This could be the case that in a planted tank with a very high turn over via filtration and powerheads, the CO2 distribution is sufficient for plants to not go in search of it. This is speculation, of course, but Hemianthus micranthemoides is one of those plants that I was convinced needed high light for it to carpet and not grow leggy. Like most of us, I presumed leggy HC meant it was growing up to the higher light levels. It turns out that, given “sufficient” light as opposed to “high light”, good levels of CO2 directed at the HC will get it carpeting nice and low. What constitutes sufficient light is down to a little experience but, ultimately, 2 x T8 tubes the length of a tank will be sufficient. Nanos are a law unto themselves, and need proportionally. more light.
If you stick the latter in a tank with poor lighting, say, 1 watt per gallon, it won't grow. CO2 will never be the limiting factor because photosynthesis isn't progressing fast enough to use up the CO2 that's already there in the water.
The WPG rule is very vague at best, but 1WPG over a reasonable sized tank such as a 120l should see Rotala sp getting by. Adding CO2 will see an increase in growth in most low light cases. As long as there is photosynthesis CO2 will improve growth proportionally.
Surely that's the case? Or am I wrong? If I add CO2 to a tank with 1 watt per gallon, can I grow Rotala, Samolus, and all those other plants adapted to strong light conditions in the wild?
I am not sure how adapted to high light these plants are. I use more than 1WPG, but my 240l has 1.2WPG and pressurised CO2, and the three Rotala sp in there have very healthy, and relatively rapid growth. The only really accurate method of determining the minimum light threshold of any given plant is to use a PAR meter, but that is out of reach for most of us. I know of one or two people measuring PAR at various depths in aquariums, and they have come to the conclusion that it is non growth limiting CO2 that is the real player, provided there is enough light, of course. CO2 injection can increase growth rates by up to x 10.
A few people, myself included, have been dragged kicking and screaming from their high light tanks, but it is CO2 that shapes our planted the most, once there is enough light. It is in the US where there appears to be the greatest reluctance to move away from high light. Even Takashi Amano`s tanks have been shown to be lower light, with a short mid day burst of light to promote faster growth.
The big deal is that we don`t need as much light as we thought, which means we can buy cheaper luminaries and use less electricity.
Dave.